In the world of Smash, challengers from various worlds come to compete in fights to see who is the best fighter. One day, as the challengers head back to their home worlds, the One Winged Angel, Sephiroth, destroys Smash as we know it. So Link, Samus, Sonic, Dark Pit, and Greninja must find a way to find the Specials and restore Smash to how it once was.